'...species react to wet, mild winters with increased activity, and can relocate to the soil surface, including euedaphic species (sometimes observed in Aporrectodea caliginosa).'
Edaphic means "of the soil", Eu- prefix means "true", so a euedaphic species is one that is native to the soil (lives underground). As opposed to epedaphic, which means above the soil (lives on the surface).
